The DSCF has columnar organization, that is, all the neurons which are perpendicular to the surface of the cortex are tuned to the same exact frequency and amplitude, whereas all the neurons which are tangential to the surface have gradually changing frequency and amplitude (Suga, 1990). ... Delays are caused by inhibitory neurons in the medial geniculate body as well as by delay lines in the inferior colliculus (Suga, 1990). ...